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The , which relies on a wide network of sources inside , said the new figure included almost 117,000 civilians, among them more than 22,000 children.

The Observatory's previous tally was issued in January and stood at more than 380,000.

The fighting, which erupted in 2011 after the brutal repression of anti-government protests, has largely abated this year as a ceasefire held in northwestern Syria and attention turned to containing the coronavirus pandemic.

The latest toll included more than 130,500 pro-government fighters, among them foreigners.

More than half of those were Syrian soldiers, while 1,703 were from the Lebanese Shiite group Hezbollah whose members have been fighting in Syria since 2013.

It has also killed more than 67,500 jihadists, mainly from the Islamic State group and Hayat Tahrir al-Sham (HTS), a group dominated by ex-members of Syria's former Al-Qaeda affiliate.

More than 12,500 Kurdish-led Syrian Democratic Forces fighters have been killed in battles against IS and Turkish forces.

The Observatory's toll does not include some 88,000 people it says died of torture in government-run prisons, nor thousands who were abducted during the conflict and are still missing.

After turning the tide of the war with Russian and Iranian support since 2015, the Syrian government now controls around two-thirds of the country.

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Among the regions still beyond its reach are the last rebel enclave of Idlib in the northwest, Turkish-held areas along the northern border, and northeastern parts of the country held by US-backed Kurdish forces.

The war has forced more than half the country's pre-war population to flee their homes.

Some 6.7 million Syrians remained displaced inside the country, while 5.5 million are registered as refugees abroad, according to the United Nations.

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