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Syria's military and Russia carried out more than 100 air raids in Idlib, Hama and Aleppo provinces, the Syrian Observatory for Human Rights said.
Rifaat al-Assad, dubbed the 'Butcher of Hama' for allegedly commanding troops that put down an uprising in Syria in 1982, has been under investigation in France since 2014.
Syrian rebels arrested 55 people in Hama and Idlib province over suspected collaboration with the regime.
Dr. Aziz Esber, director of the Center for Scientific Research in Masyaf was killed in an attack in Hama province near the town of Masyaf.
Scores of Syrian Faylaq al-Rahman rebels are arriving in Hama, after being evacuated from Eastern Ghouta's Arbin, leaving only one town under rebel control in the besieged Damascus enclave.
The Islamic State group previously held large swathes of northern and central Syria, but has been fully ousted from Idlib, Hama and Aleppo provinces, a monitor has said.
Syrian regime forces, backed by Russian airstrikes, have captured nearly 100 villages from rebels since late October in Hama and Idlib provinces.
Two pilots were killed and a third person injured after a Russian military helicopter crashed in Syria while flying to Hama.
The refugees and fighters from al-Qaeda's former Syrian branch arrived in bus convoys into the rebel-held area of Al-Saan in the central Syrian Hama province.
Activists say the Syrian regime and Russian warplanes targeted the Idlib and Hama provinces with incendiary bombs, including thermite.